1915683 - Errors 2601, 3461, 3451, 3454 when online database after upgrade from 12.5.1 to 15.5 ESD #5 - SAP ASE

Upgrade from ASE 12.5.1 to ASE 15.5 ESD#5, getting errors during online of a database:

Initialization of sysusers in database <database> changed the system role name from '<user>' to 'dtm_tm_role'.
 
Msg 2601, Level 14, State 6:
Server '<server>', Line 1:
Attempt to insert duplicate key row in object 'sysusers' with unique index 'ncsysusers1'
Command has been aborted.
(0 rows affected)
 
Msg 3461, Level 20, State 1:
Server '<server>', Line 1:
Database '<database>': upgrade could not install required upgrade item '90'.
Please refer to previous error messages to determine the problem. Fix the problem, then try again.
 
Msg 3451, Level 20, State 1:
Server '<server>', Line 1:
Database '<database>': upgrade has failed for this database.  Please refer to previous messages for the cause of the failure, correct the problem and try again.
 
Msg 3454, Level 20, State 1:
Server '<server>', Line 1:
Database '<database>': ASE could not completely upgrade this database; upgrade item 90 could not be installed.
ASE could not bring database '<database>' online.

• SAP Adaptive Server Enterprise (ASE) 15.0
• Sybase Adaptive Server Enterprise (ASE) 15.5

CR569005,  CR#569005,  569005, CR614043, CR#614043, 614043, syssrvroles, syssrvroles,permission, sqlupgrade, sqlupgraderes, DTM_TM_ROLE, HA_ROLE, MON_ROLE, MESSAGING_ROLE, JS_ADMIN_ROLE, traceflag, 990 , KBA , BC-SYB-ASE , Sybase ASE Database Platform (non Business Suite) , Bug Filed