3241137 - Failed to execute ..., because this is a read-only connection - SAP ASE

After upgrading to ASE 16 SP04 GA and higher, queries are raising following (similar) errors: 

• java.sql.SQLException: Failed to execute COMMIT TRANSACTION, because this is a read-only connection. Read-only connections can only execute SELECT statements. Use 'set readonly off' to execute COMMIT TRANSACTION.
• com.sybase.jdbc3.jdbc.SybSQLException: Failed to execute ROLLBACK TRANSACTION, because this is a read-only connection. Read-only connections can only execute SELECT statements. Use 'set readonly off' to execute ROLLBACK TRANSACTION. 
• com.sybase.jdbc4.jdbc.SybSQLException: Failed to execute DROP PROCEDURE, because this is a read-only connection. Read-only connections can only execute SELECT statements. Use 'set readonly off' to execute DROP PROCEDURE. 

SAP Adaptive Server Enterprise (ASE) 16.0 SP04 GA and higher

Failed to execute, COMMIT TRANSACTION, ROLLBACK TRANSACTION, DROP PROCEDURE, read-only connection, read-only, read only, set readonly off, set readonly, readonly off, CR827215, CR#827215, 827215 , KBA , BC-SYB-ASE , Sybase ASE Database Platform (non Business Suite) , Known Error